"The competition of races and species, observes Mr. Darwin, is always most severe between those which are most closely allied and which fill nearly the same place in the economy of nature. Hence, when the conditions of existence are modified, the original stock runs great risk of being superseded by some one of its modified offshoots. The new race or species may not be absolutely superior in the sum of its powers and endowment to the parent stock, and may even be more simple in structure and of a lower grade of intelligence, as well as of organisation, provided, on the whole, it happens to have some slight advantage over its rivals. Progression, therefore, is not a necessary accompaniment of variation and natural selection, though, when a higher organisation happens to be coincident with superior fitness to new conditions, the new species will have greater power and a greater chance of permanently maintaining and extending its ground."

“Lamarck, when speculating on the origin of the long neck of the giraffe, imagined that quadruped to have stretched himself up in order to reach the boughs of lofty trees, until by continued efforts, and longing to reach higher, he obtained an elongated neck. Mr. Darwin and Mr. Wallace simply suppose that, in a season of scarcity, a longer-necked variety, having the advantage in this respect over most of the herd, as being able to browse on foliage out of their reach, survived them, and transmitted its peculiarity of cervical conformation to its successors.”

“The form of a coast, the configuration of the interior of a country, the existence and extent of lakes, valleys, and mountains, can often be traced to the former prevalence of earthquakes and volcanoes in regions which have long been undisturbed. To these remote convulsions the present fertility of some districts the sterile character of others, the elevation of land above the sea, the climate, and various peculiarities, may be distinctly referred.”

“The era, it is true, of the expected millennium had passed away; and for five hundred years after the fatal hour, when the annihilation of the planet had been looked for, the monks remained in undisturbed enjoyment of rich grants of land bequeathed to them by pious donors, who, in the preamble of deeds beginning 'appropinquante mundi termino'-'appropinquante magno judicii die,' left lasting monuments of the popular delusion.”

“What Lamarck then foretold has come to pass; the more new forms have been multiplied, the less are we able to decide what we mean by a variety, and what by a species. In fact, zoologists and botanists are not only more at a loss than ever how to define a species, but even to determine whether it has any real existence in nature, or is a mere abstraction of the human intellect, some contending that it is constant within certain narrow and impassable limits of variability, others that it is capable of indefinite and endless modification.”

“The theory of the origin of new species by variation will also explain why a species which has once died out never reappears, and why the fossil fauna and flora recede farther and farther from the living type in proportion as we trace it back to remoter ages. It would also account for the fact, that when we have to intercalate a new set of fossiliferous strata between two groups previously known, the newly discovered fossils serve to fill up gaps between specific or generic types previously familiar to us, supplying often the missing links of the chain, which, if transmutation is accepted, must once have been continuous.”

“We may understand why the species of the same genus, or genera of the same family, resemble each other more nearly in their embryonic than in their more fully developed state, or how it is that in the eyes of most naturalists the structure of the embryo is even more important in classification than that of the adult, 'for the embryo is the animal in its less modified state, and in so far it reveals the structure of its progenitor. In two groups of animals, however much they may at present differ from each other in structure and habits, if they pass through the same or similar embryonic stages, we may feel assured that they have both descended from the same or nearly similar parents, and are therefore in that degree closely related. Thus community in embryonic structure reveals community of descent, however much the structure of the adult may have been modified.”